The protected area covers some 685 km², consisting of a central uninhabited zone comprising seven valleys - Roya, Bévéra, Vésubie, Tinée, Haut Var/Cians (in the Alpes-Maritimes) plus Verdon and Ubaye (in the Alpes-de-Haute-Provence) - and a peripheral zone comprising 28 villages. Many of them are perched villages,such as Belvédère at the entrance to the spectacular Gordolasque valley, concealing great architectural riches (numerous churches decorated with murals and altar pieces by primitive Niçois painters).
More than 150 rural sites are located within the Park.
Around Mont Bégo there are petroglyphs pecked out on schist and granite faces. They have been dated from the late Neolithic and Bronze Ages.